White Shark, Carcharodon carcharias (Linnaeus 1758)
A White Shark, Carcharodon carcharias, at Isla Guadalupe, Mexico. Source: sharkdiver.com / Wikimedia Commons. License: Public Domain
A large powerful shark with a very high first dorsal fin, a crescent-shaped tail, large triangular serrate teeth, and minute anal and second dorsal fins. Blue-grey to greyish-brown above, abruptly changing to a white underside.
White Sharks are warm-blooded. They have a special heat-exchange circulatory system that allows them to maintain body temperatures up to 14C higher than the surrounding seawater. Adults usually feed on seals, penguins, fishes and seabirds, and this dangerous apex predator is responsible for many fatal attacks on humans.
White Sharks are protected in all Australian waters, including territorial waters - and in many other parts of the world.
Video of tagging a juvenile white shark with an internal acoustic tag at Port Stephens, New South Wales.
See what's inside a White Shark that died after being caught in a net off South Africa on The Evolution Documentary YouTube Channel Inside Nature's Giants 5/18 Great White Shark
White Shark, Carcharodon carcharias (Linnaeus 1758)
Occurs worldwide, mostly in temperate waters, but also ventures into tropical and subtropical waters. In Australia, White Sharks are known from about Yeppoon on the Queensland coast (and even further north in the Coral Sea), around southern Australia to about Onslow in north-western Western Australia.
White sharks are found from inshore waters around rocky reefs, surf beaches and shallow coastal bays, to waters on the outer continental shelf and slope. Although they most commonly live in depths above 100 m, both juveniles and adults are known to dive to at least 1000 m.
Individuals may also undertake long open ocean journeys, including crossing the Indian Ocean, swimming coast-to-coast from South Africa to Australia and back.
Research indicates that at the global scale, three distinct geographical White Shark clades (Andreotti et al. 2015). In Australia, there appear to be two White Shark populations - an eastern and a western population. Tagging studies indicate that White Sharks in the eastern population may breed in Bass Strait, and that Corner Inlet, (Victoria) and the Port Stephens region (New South Wales) are important nursery areas for juveniles.
Vertebrae: 172-187, (precaudal 100-108); Jaw teeth (upper) 13:13; Jaw teeth (lower) 12:12
|To a maximum length of at least 6.4 metres.|
Greyish to greyish-brown above, white below, with a distinct separation between the dark and light pigment.
Voracious carnivores. Adult White sharks are opportunistic feeders, and prey on a range of vertebrates. Prey items include bottom-dwelling and schooling fishes, large species including crustaceans, cephalopod molluscs; broadbill swordfish and bluefin tuna, sharks, rays and chimaeras; turtles; occasionally sea birds such as cormorants and penguins; seals, sea lions, and small whales. They also feed on dead baleen whales.
Malcolm et al. (2001) reported that the smallest white shark known from Australian waters to contain seal remains was a 2.7m individual. However, the stomach of a 2.4 m juvenile male caught off Lakes Entrance, Victoria, contained a seal skull and several bony fishes.
White sharks also feed on sea birds, turtles and ocean sunfish (Mola), and both juveniles and adults scavenge on floating whale carcasses. Bruce & Stevens (2004) report that white sharks are particularly active around whale strandings, presumably because they can smell whale oil and rotting flesh in the water.
White sharks exhibit viviparous (live bearers) and oophagous reproduction (embryos feed on developing eggs in the uterus).
Life history characteristics:
|White Sharks are protected in Australia, including in all state and territorial waters.|
IUCN Red List: Vulnerable
EPBC Act: Listed as Vulnerable and Migratory under the Environment Protection and Biodiversity Conservation Act 1999.
New South Wales: Listed as Vulnerable under Section 5, Part 1, Fisheries Management Act 1994.
Queensland: Protected under Schedule 78 (1), Fisheries Act 1994.
South Australia: Protected under Schedule 42, Fisheries Act 1982.
Tasmania: Protected under Schedule 135(2), Threatened Species Protection Act 1995, Section 135(2), Living Marine Resources Management Act 1995 and declared vulnerable under the Fisheries (General and Fees) Regulations 1996.
Victoria: Protected under Schedule 71, Fisheries Act 1995.
Western Australia: Listed as rare or likely to become extinct under Schedule 5 of the Wildlife Conservation Act 1950 and protected under Schedule 46 of the Fisheries Resources Management Act 1994. In response to a number of shark attacks in WA, in December 2013, the Western Australian Government announced a number of measures to combat shark risks.
White Sharks are protected in all Australian maritime waters, and in many other parts of the world, including in New Zealand.
Commercial tourist operations for the viewing of White Sharks were established in Spencer Gulf, South Australia in the 1970's, however, the White Shark cage dive industry has increased rapidly in recent years.
Charter companies operating from Port Lincoln, provide tourists with opportunites to view sharks at the Neptune Islands, 60-70 km south of Port Lincoln. These operations are very successful and provide world-class shark cage-diving experiences.
The Neptune Islands have the largest seal populations in Australia, and vessel operators attract White Sharks with a berley/chum mix of minced tuna and tuna oil. Baits of tuna pieces and entrails are also tied to the vessels.
Recent studies have shown that there have been some changes to white shark behaviour and residency at the North Neptune Islands since 2007 when the industry increased the number and regularity of cage diving activities (Bruce & Bradford 2013). As these changes may lead to long-term localised changes in the behaviour of White Sharks, cage diving tourism can only operate 5 out of 7 days per week.
Carcharodon is from the Greek carcharos (ragged) and odon (tooth), in reference to the triangular serrated teeth of this species.
Squalus carcharias Linnaeus, 1758, Systema Naturae: 235. Type locality: Europe.
Bray, D.J. 2017
Read the excellent 2009 White Shark Issues Paper and the 2013 Recovery Plan for the White Shark (Carcharodon carcharias) by the Department of the Environment, Water, Heritage and the Arts.
White Shark, Carcharodon carcharias (Linnaeus 1758)
Andreotti, S., von der Heyden, S., Henriques, R., Rutzen, M., Meÿer, M., Oosthuizen, H. & Matthee, C.A. 2015. New insights into the evolutionary history of white sharks, Carcharodon carcharias. Journal of Biogeography. doi: 10.1111/jbi.12641 Abstract
Blower, D.C., Pandolfi, J.M., Bruce, B.D., Gomez-Cabrera, M.C. & Ovenden, J.R. 2012. Population genetics of Australian white sharks reveals fine-scale spatial structure, transoceanic dispersal events and low effective population sizes. Marine Ecology Progress Series 455: 229–244.
Bonfil, R., M.P. Francis, C. Duffy, M.J. Manning & S.M. O’Brien. 2010. Large-scale tropical movements and diving behaviour of white sharks (Carcharodon carcharias) tagged off New Zealand. Aquatic Biology 8: 115–123.
Bonfil, R., M. Meer, M.C. Scholl, R. Johnson, S. O’Brien, H. Oosthuizen, S. Swanson, D. Kotze & M. Paterson. 2005. Transoceanic migration, spatial dynamics, and population linkages of white sharks. Science 310(5745): 100-103. DOI: 10.1126/science.1114898.full Open access
Boustany, A.M., S.F. Davis, P. Pyle, S.D. Anderson, B.J. Le Boeuf & B.A. Block. 2002. Expanded niche for white sharks. Nature 415: 35–36.
Bradford, R. & R. Robbins. 2013. A rapid assessment technique to assist management of the White Shark (Carcharodon carcharias) cage dive industry, South Australia. The Open Fish Science Journal 6: 13-18.
Briscoe, D.A., Stow, A. & Hussey, S. 2006. Trial DNA Testing of shark products imported into and exported from Australia to detect the presence of Great White Shark. Report to the Department of the Environment and Heritage. May 2005.
Bruce, B.D. 1992. Preliminary observations on the biology of the White Shark, Carcharodon carcharias, in South Australian waters. Australian Journal of Marine and Freshwater Research 43: 1-11.
Bruce, B.D. 1995. The Protection of White Shark. A research perspective. Southern Fisheries 3(2): 11-15.
Bruce, B.D. 2006. Coastal migrations of white sharks in Australian waters: Evidence for site fidelity, temporary residency and common pathways (pp 67-77). In DPI NSW. Report from the Scientific Shark Protection Summit, 10 April 2006, 104 pp. PDF
Bruce, B.D. 2008. The Biology and Ecology of the White Shark, Carcharodon carcharias. In: Camhi, M.D, E.K. Pikitch & E.A Babcock (eds). Sharks of the Open Ocean. Pp 69-76. Oxford, UK: Blackwell Publishing.
Bruce, B.D & Bradford, R.W. 2008. Spatial dynamics & habitat preferences of juvenile white sharks: identifying critical habitat and options for monitoring recruitment. Final Report to the Department of the Environment, Water, Heritage and the Arts - Marine Species Recovery Program. Hobart: CSIRO.
Bruce, B. & Bradford, R.W. 2011. The effects of burleying on the distribution and behaviour of white sharks, Carcharodon carcharias, at the Neptune Islands, South Australia: final report. Department of Environment and Natural Resources, South Australia.
Bruce, B.D. & Bradford, R.W. 2012. Habitat use and spatial dynamics of juvenile white sharks, Carcharodon carcharias, in Eastern Australia. In Domeier, M. (ed.) Global perspectives on the biology and life history of the white shark. CRC Press, Boca Raton, FL.
Bruce, B.D. & Bradford, R.W. 2013. The effects of shark-cage diving operations on the behavior and movements of white sharks, Carcharodon carcharias, at the Neptune Islands, South Australia. Mar Biol ; DOI: 10.1007/s00227-012-2142-z
Bruce, B.D. & Bradford, R.W. 2015. Segregation or aggregation? Sex-specific patterns in the seasonal occurrence of white sharks Carcharodon carcharias, at the Neptune Islands, South Australia. Journal of Fish Biology 87: 1355-1370.
Bruce. B.D., Malcolm H. & Stevens J.D. 2001. A Review of the Biology and Status of White Sharks in Australian Waters. CSIRO Marine Research, Hobart.
Bruce, B.D., J.D. Stevens & R.W. Bradford. 2005. Site Fidelity, Residence Times and Home Range Patterns of White Sharks around Pinniped Colonies. [Online]. Final Report to the Australian Government Department of the Environment and Heritage, Canberra.
Bruce, B.D., J.D. Stevens & R.W. Bradford. 2005. Identifying movements and habitats of white sharks and grey nurse sharks. [Online]. Available from: http://www.environment.gov.au/coasts/publications/white-grey-nurse-habitats/index.html.
Bruce, B.D., Stevens, J.D. & Malcolm, H. 2006. Movements and swimming behaviour of white sharks (Carcharodon carcharias) in Australian waters. Marine Biology 150: 161-172.
Cailliet, G.M., Natanson, L.J., Weldon, B.A. & Ebert, D.A. 1985. Preliminary studies on the age and growth of the white shark Carcharodon carcharias, using vertebral bands. Southern California Academy of Sciences, Memoirs 9: 49-60.
Carey, F.G., Kanwisher, J.W., Brazier, O., Gabrielson, G., Casey, J.G. & Pratt Jr, H.L. 1982. Temperature and activities of a white shark, Carcharodon carcharias. Copeia 1982: 254-260.
Castro J [Domeier M]. 2012. A Summary of Observations on the Maximum Size Attained by the White Shark, Carcharodon carcharias. Global Perspectives on the Biology and Life History of the White Shark. Boca Raton, FL: CRC Press. 85-90 p.
CITES (2004). Convention of International Trade in Endangered Species of Wild Fauna and Flora - Appendix II Listing of the White Shark (revision 1).
Compagno, L.J.V. 1984. FAO Species Catalogue. Sharks of the World. An annotated and illustrated catalogue of shark species known to date. Hexanchiformes to Lamniformes. FAO Fisheries Synopsis No. 125. Rome : FAO Vol. 4(1) pp. 1-249.
Compagno, L.J.V. 1998. Families Pseudocarchariidae, Alopiidae, Lamnidae. pp. 1268-1278 in Carpenter, K.E. & Niem, V.H. (eds). The Living Marine Resources of the Western Central Pacific. FAO Species Identification Guide for Fisheries Purposes. Rome : FAO Vol. 2 687-1396 pp.
Compagno, L.J.V. 2001. Sharks of the World. An annotated and illustrated catalogue of shark species known to date. Bullhead, mackerel and carpet sharks (Heterodontiformes, Lamniformes and Orectolobiformes). Rome : FAO, FAO Species Catalogue for Fisheries Purposes No. 1 Vol. 2 269 pp.
Compagno, L.J.V., Dando, M. & Fowler, S. 2005. A Field Guide to the Sharks of the World. London : Collins 368 pp.
Compagno, L.J.V., Marks, M.A. & Fergusson, I.K. 1997. Threatened fishes of the world: Carcharodon carcharias (Linnaeus, 1758) (Lamnidae). Environmental Biology of Fishes 50: 61-62.
Del Raye, G., S.J. Jorgensen, K. Krumhans, J.M. Ezcurra & B.A. Block. 2013. Travelling light: white sharks (Carcharodon carcharias) rely on body lipid stores to power ocean-basin scale migration. Proc. R. Soc. B 280: 1471-2954. PDF
Department of the Enviroment, Water, Heritage and the Arts (DEWHA). 2009. White Shark Issues Paper. [Online]. Canberra, ACT: Australian Government.
Department of the Environment, Water, Heritage and the Arts (DEWHA). 2010. Draft Recovery Plan for the White Shark (Carcharodon carcharias). [Online]. Canberra, ACT: Australian Government.
Dicken, M.L. 2008. First observations of young of the year and juvenile great white sharks (Carcharodon carcharias) scavenging from a whale carcass. Marine and Freshwater Research 59: 596-602.
Domeier ML (2012) A new life-history hypothesis for white sharks, Carcharodon carcharias, in the northeast Paciﬁc. Pp 199–223, in: Domeier M (ed) Global perspectives on the biology and life history of the Great White Shark. CRC Press, Boca Raton.
Ellis, R. & J.E. McCosker. 1991. Great White Shark. Stanford University Press. 270 pp.
Estrada, J.A., Rice, A.N., Natanson, L J. & Skomal, G.B. 2006. Use of isotopic analysis of vertebrae in reconstructing ontogenetic feeding ecology in white sharks. Ecology 87: 829-834.
Fergusson, I., Compagno, L.J.V. & Marks, M.A. 2000. Predation by white sharks Carcharodon carcharias upon chelonians, with new records from the Mediterranean Sea and a first record of the ocean sunfish Mola mola as stomach contents. Environmental Biology of Fishes 58: 447-453.
Fergusson, I., Compagno, L.J.V. & Marks, M. 2009. Carcharodon carcharias. The IUCN Red List of Threatened Species 2009: e.T3855A10133872. http://dx.doi.org/10.2305/IUCN.UK.2009-2.RLTS.T3855A10133872.en. Downloaded on 02 October 2016.
Fitzpatrick, B., M. Meekan & A. Richards. 2006. Shark attacks on a whale shark (Rhincodon typus) at Ningaloo Reef, Western Australia. Bull. Mar. Sci. 78: 397-402.
Fowler, S.L., Cavanagh, R.D., Camhi, M., Burgess, G.H., Cailliet, G.M., Fordham, S.V., Simpfendorfer, C.A. and Musick, J.A. (comps and eds) 2005. Sharks, Rays and Chimaeras: The Status of the Chondrichthyan Fishes. Status Survey. pp. x + 461. IUCN/SSC Shark Specialist Group, IUCN, Gland, Switzerland and Cambridge, UK.
Francis, M.P. 1996. Observations on a pregnant White Shark with a review of reproductive biology (pp 157-172). In: Klimley, A.P. & D.G Ainley (eds). Great White Sharks. The biology of Carcharodon carcharias. Academic Press, San Diego.
Goldman, K.J. 1997. Regulation of body temperature in the white shark, Carcharodon carcharias. Journal of Comparative Physiology B. Biochemical Systemic and Environmental Physiology. 167: 423-429.
Goldman, K.J., Anderson, S.D., McCosker, J.E. & Klimley, A.P. 1996. Temperature, swimming depth, and movements of a White Shark at the South Farallon Islands, California. In: Klimley, A.P. & D.G Ainley (eds). Great White Sharks. The biology of Carcharodon carcharias. Academic Press, San Diego.
Gubili C, Duffy CAJ, Cliff G, Wintner SP, Shivji M, Chapman D, Bruce BD, Martin AP, Sims DW [Domeier M]. 2012. Application of molecular genetics for conservation of the White Shark, Carcharodon carcharias, L. 1758. Global Perspectives of the Life History of the White Shark. Boca Raton, FL: CRC Press. 357-380 p.
Hammerschlag N, Gallagher AJ, Wester J, Luo J, Ault JS (2012) Don’t bite the hand that feeds: assessing the ecological impacts of provisioning ecotourism on an apex marine predator. Funct Ecol. doi:10.1111/j.1365-2435.2012.01973.x
Hutchins, B. & R. Swainston. 1986. Sea Fishes of Southern Australia. Complete Field Guide for Anglers and Divers. Swainston Publishing. 180 pp.
Johnson, J.W. 2010. Fishes of the Moreton Bay Marine Park and adjacent continental shelf waters, Queensland, Australia. 299-353 in Davie, P.J.F. & Phillips, J.A. Proceedings of the Thirteenth International Marine Biological Workshop, The Marine Fauna and Flora of Moreton Bay. Memoirs of the Queensland Museum 54(3) 
Jorgensen, S.J., C.A. Reeb, T.K. Chapple, S. Anderson, C. Perle, S.R. Van Sommeran, C. Fritz-Cope, A.C. Brown, A.P. Klimley & B.A. Block. 2010. Philopatry and migration of Pacific white sharks. Proc. R. Soc. B 277: 679–688. doi:10.1098/rspb.2009.1155 PDF
Klimley, A.P. 1994. The predatory behaviour of the White Shark. American Scientist 82: 122-133.
Klimley, A.P. & D.G Ainley (eds). 1996. Great White Sharks. The biology of Carcharodon carcharias. Academic Press, San Diego. 517pp.
Klimley, A.P., Le Boeuf, B. J., Cantara, K. M., Richert, J. E., Davis, S. F., van Sommeran, S. & Kelly, J.T. 2001. The hunting strategy of white sharks (Carcharodon carcharias) near a seal colony. Marine Biology 138: 617-636.
Kuiter, R.H. 1993. Coastal Fishes of South-eastern Australia. Bathurst : Crawford House Press 437 pp.
Lack M, Sant G. 2011. The Future of Sharks: A Review of Action and Inaction. Cambridge, U.K. and Washington D.C. TRAFFIC International and the Pew Environment Group. 45 p.
Laroche RK, Kock AA, Dill LM, Oosthuizen WH. 2007. Effects of provisioning ecotourism activity on the behaviour of white sharks, Carcharodon carcharias. Mar Ecol Prog Ser 338: 199-209.
Last, P.R. & Stevens, J.D. 1994. Sharks and Rays of Australia. Canberra : CSIRO Australia 513 pp. 84 pls
Last, P.R. & Stevens, J.D. 2009. Sharks and Rays of Australia. Vol 2. Collingwood : CSIRO Publishing Australia, 656pp.
Lingham-Soliar T. 2005. Caudal fin allometry in the white shark Carcharodon carcharias: implications for locomotory performance and ecology. Die Naturwiss. 92: 231– 236. (doi:10.1007/s00114-005-0614-4)
Linnaeus, C. 1758. Systema Naturae per Regna tria Naturae, secundem Classes, Ordines, Genera, Species, cum Characteribus, Differentis, Synonymis, Locis. Tom.1, Editio decima, reformata. Holmiae : Laurentii Salvii 824 pp.
Macbeth, W.G., Vandenberg, M. & Graham, K.J. 2008. Identifying Sharks and Rays; a Guide for Commercial Fishers. Sydney : New South Wales Department of Primary Industry 71 pp.
Malcolm, H., B.D. Bruce & J. Stevens. 2001. A Review of the Biology and Status of White Sharks in Australian Waters. Environment Australia, Canberra. 113 pp.
Mcauley, R.D., Bruce, B.D., Keay, I.S., Mountford, S., Pinnell, T. & Whoriskey, F.G. 2017. Broad-scale coastal movements of white sharks off Western Australia described by passive acoustic telemetry data. Marine and Freshwater Research. 68(8) 1518-1531. https://doi.org/10.1071/MF16222 Abstract
McPherson, J.M & R.A Myers. 2009. How to infer population trends in sparse data: examples with opportunistic sighting records for great white sharks. Diversity and Distributions. 15: 880-890.
Pardini, A.T., Jones, C.S., Noble, L.R., Kreiser, B., Malcolm, H., Bruce, B.D., Stevens, J.D., Cliff, G., Scholl, M.C., Francis, M., Duffy, C. & Martin, A.P. 2001. Philopatric females and roving male great white sharks. Nature 412: 139–140.
Pardini, A.T., Jones, C.S., Noble, L.R., Kreiser, B., Malcolm, H., Bruce, B.D., Stevens, J.D., Cliff, G., Scholl, M.C., Francis, M., Duffy, C. & Martin, A.P. 2001. Sex biased dispersal of Great White Sharks. Nature 412: 139-140.
Pepperell, J. 1992. Trends in the distribution, species composition and size of sharks caught by gamefish anglers off south-eastern Australia, 1961-1990. In: J. Pepperell (ed.), Sharks: Biology and Fisheries. Australian Journal of Marine and Freshwater Research, pp. 213-225.
Pogonoski, J.J., Pollard, D.A.& J.R. Paxton. 2002. Conservation Overview and Action Plan for Australian Threatened and Potentially Threatened Marine and Estuarine Fishes. Environment Australia, ISBN 0 642 54786 6.
Pepperell, J. 2010. Fishes of the Open Ocean a Natural History & Illustrated Guide. Sydney : University of New South Wales Press Ltd 266 pp.
Pratt, Jr. H.L. 1996. Reproduction in the male White Shark. In: Klimley, A.P. & D.G. Ainley (eds). Great White Sharks: The Biology of Carcharodon carcharia. Academic Press, San Diego.
Randall, J.E. 1973. Size of the great white shark (Carcharodon). Science (Washington, D.C.) 181(4095): 169-170
Reid DD, Krogh M. 1992. Assessment of catches from Protective Shark meshing off New South Wales beaches between 1950 and 1990. Australian Journal of Marine and Freshwater Research 43(1): 283-296.
Robbins, R.L. 2007. Environmental variables affecting the sexual segregation of great white sharks Carcharodon carcharias at the Neptune Islands South Australia. Journal of Fish Biology 70: 1350-1364.
Robbins, R.L. & D.J. Booth. 2012. Seasonal sexual and size segregation of white sharks, Carcharodon carcharias, at the Neptune Islands, South Australia. Pp. 287-99, in Domeier, M.L. (ed.) Global perspectives on the biology and life history of the white shark. Boca Raton, Fl: CRC Press.
Robbins RL, Enarson M, Bradford RW, Robbins WD, Fox AG. 2015. Residency and Local Connectivity of White Sharks at Liguanea Island: A Second Aggregation Site in South Australia? The Open Fish Science Journal, 2015, 8: 23-29. [DOI: 10.2174/1874401X01508010023] PDF Open Access
Saïdi, B., Bradaï, M.N., Bouaïn, A., Guélorget, O. & C. Capapé. 2005. Capture of a pregnant female white shark, Carcharodon carcharias (Lamnidae) in the Gulf of Gabés (southern Tunisia, central Mediterranean) with comments on oophagy in sharks. Cybium 29(3): 303-307.
Sato, K., Nakamura, M., Tomita, T., Toda, M., Miyamoto, K. & Nozu, R. 2016, How great white sharks nourish their embryos to a large size: evidence of lipid histotrophy in lamnoid shark reproduction. Biology Open 5: 1211-1215; doi: 10.1242/bio.017939 Open access
Semmens, J.M., Payne, N.L., Huveneers, C., Sims, D.W. & Bruce, B.D. 2013. Feeding requirements of white sharks may be higher than originally thought. Scientific Reports 3: 1471
Shivji, M. S., D.D. Chapman, E.K. Pikitch, & P.W. Raymond (2005). Genetic profiling illegal international trade in fins of the great white shark, Carcharodon carcharias. Conservation Genetics 6: 1035-1039.
Sims DW, Humphries NE, Bradford RW, Bruce BD (2012) Levy ﬂight and brownian search patterns of white sharks in response to different prey ﬁelds. J Anim Ecol 81: 432–442.
Stevens, J.D. 1994. Family Lamnidae. pp. 141-143 figs 112-114 in Gomon, M.F., Glover, C.J.M. & Kuiter, R.H (eds). The Fishes of Australia's South Coast. Adelaide : State Printer 992 pp. 810 figs
Strong, W.R., Nelson, D.R., Bruce, B.D. and Murphy, R.C. 1996. Population dynamics of White Sharks in Spencer Gulf, South Australia (pp. 401-414). In: Klimley, A.P. & D.G. Ainley, (eds). Great White Sharks - The Biology of Carcharodon carcharias. Academic Press, San Diego.
Strong, W.R., Murphy, R.C., Bruce, B.D. and Nelson, D.R. 1992, Movements and associated observations of bait-attracted White Sharks, Carcharodon carcharias: A preliminary report. Australian Journal of Marine and Freshwater Research 43: 13-20.
Tricas, T.C. & J.E. McCosker. 1984. Predatory behaviour of the white shark (Carcharodon carcharias) with notes on its biology. roceedings of the California Academy of Sciences. 43: 221-238.
Uchida, S., Toda, M., Teshima, K. and Yano, K. 1996. Pregnant White Shark and full-term embryos from Japan (pp 139-155). In: Klimley, A.P. & D.G. Ainley, (eds). Great White Sharks - The Biology of Carcharodon carcharias. Academic Press, San Diego.
Walker, T. 1999. Protection for Great Whites. Sport Fishing Issue 5.
White, W. 2008. Shark Families Heterodontidae to Pristiophoridae. pp. 32-100 in Gomon. M.F., Bray, D.J. & Kuiter, R.H (eds). Fishes of Australia's Southern Coast. Sydney : Reed New Holland 928 pp.
White Sharks. Filling the gaps in our knowledge. CSIRO fact sheet 35, May 1999, Pp. 2. White Shark Research. Neale the white shark. CSIRO Australia.