White Shark, Carcharodon carcharias (Linnaeus 1758)
A White Shark, Carcharodon carcharias, at Isla Guadalupe, Mexico. Source: sharkdiver.com / Wikimedia Commons. License: Public Domain
A large powerful shark with a very high first dorsal fin, a crescent-shaped tail, large triangular serrate teeth, and minute anal and second dorsal fins. Blue-grey to greyish-brown above, abruptly changing to a white underside.
White Sharks are warm-blooded. They have a special heat-exchange circulatory system that allows them to maintain body temperatures up to 14C higher than the surrounding seawater. Adults usually feed on seals, penguins, fishes and seabirds, and this dangerous apex predator is responsible for many fatal attacks on humans.
White Sharks are protected in all Australian waters, including territorial waters - and in many other parts of the world.
Video of White Sharks at Isla Guadalupe
Video of free-diving with White Sharks
Video of tagging a juvenile white shark with an internal acoustic tag at Port Stephens, New South Wales.
See what's inside a White Shark that died after being caught in a net off South Africa on The Evolution Documentary YouTube Channel Inside Nature's Giants 5/18 Great White Shark
White Shark, Carcharodon carcharias (Linnaeus 1758)
In Australia, White Sharks are known from about Yeppoon on the Queensland coast (and even further north to Innisfail and reefs in the Coral Sea), around southern Australia to about Onslow, north-western Western Australia. The species also occurs in at Lord Howe Island and Norfolk Island in the Tasman Sea.
White sharks are found from inshore waters around rocky reefs, surf beaches and shallow coastal bays, to waters on the outer continental shelf and slope. Although most commonly found in depths above 100 m, White Sharks are known to dive to at least 1300 m.
Individuals may undertake long open ocean journeys, including crossing the Indian Ocean, swimming coast-to-coast from South Africa to Australia and back.
Research indicates that at the global scale, three distinct geographical White Shark clades (Andreotti et al. 2015). In Australia, there appear to be two White Shark populations - an eastern and a western population. Tagging studies indicate that White Sharks in the eastern population may breed in Bass Strait, and that Corner Inlet, (Victoria) and the Port Stephens region (New South Wales) are important nursery areas for juveniles.
Although mostly a solitary species, White Sharks seasonally aggregate at a number of sites worldwide to feed on calorie-rich pinnipeds (seals and sea lions) (Schilds et al. 2019).
Vertebrae: 172-187, (precaudal 100-108); Jaw teeth (upper) 13:13; Jaw teeth (lower) 12:12
|To a maximum length of at least 6.4 metres.|
Greyish to greyish-brown above, white below, with a distinct separation between the dark and light pigment.
Voracious carnivores. Adults are opportunistic feeders that prey on a range of vertebrates, including fishes, large crustaceans, cephalopod molluscs, turtles, sea birds, seals, sea lions and small whales. They also feed on dead baleen whales.
Malcolm et al. (2001) reported that the smallest white shark known from Australian waters to contain seal remains was a 2.7m individual. However, the stomach of a 2.4 m juvenile male caught off Lakes Entrance, Victoria, contained a seal skull and several bony fishes.
Juveniles feed on mid-water fishes like Australian salmon, bottom-dwelling fishes such as flounders and stingrays, and occasionally on reef-dwelling fishes.
White sharks exhibit viviparous (live bearers) and oophagous reproduction (embryos feed on developing eggs in the uterus).
Life history characteristics:
|White Sharks are protected in Australia, including in all state and territorial waters.|
IUCN Red List: Vulnerable
EPBC Act: Listed as Vulnerable and Migratory under the Environment Protection and Biodiversity Conservation Act 1999.
NSW: Listed as Vulnerable (Fisheries Management Act 1994 (New South Wales): August 2016 list) as Carcharadon carcharias
TAS: Listed as Vulnerable (Threatened Species Protection Act 1995 (Tasmania): June 2019 list) as Carcharodon carcharias
VIC: Listed as Threatened (Flora and Fauna Guarantee Act 1988 (Victoria): April 2018 list) as Carcharodon carcharias
WA: Listed as Vulnerable (Biodiversity Conservation Act 2016 (Western Australia): September 2018 list) as Carcharodon carcharias
White Sharks are protected in all Australian maritime waters, and in many other parts of the world, including in New Zealand.
Although commercial tourist operations for viewing White Sharks were established in Spencer Gulf, South Australia in the 1970's, the White Shark cage dive industry has increased rapidly in recent years.
Charter companies operating from Port Lincoln, provide tourists with opportunites to view sharks at the Neptune Islands, 60-70 km south of Port Lincoln. These operations are very successful and provide world-class shark cage-diving experiences.
The Neptune Islands have the largest seal populations in Australia, and vessel operators attract White Sharks with a berley/chum mix of minced tuna and tuna oil. Baits of tuna pieces and entrails are also tied to the vessels.
Recent studies have shown that there have been some changes to white shark behaviour and residency at the North Neptune Islands since 2007 when the industry increased the number and regularity of cage diving activities (Bruce & Bradford 2013). As these changes may lead to long-term localised changes in the behaviour of White Sharks, cage diving tourism can only operate 5 out of 7 days per week.
Carcharodon is from the Greek carcharos (ragged) and odon (tooth), in reference to the triangular serrated teeth of this species.
The specific name carcharias is from the Greek karcharias (= man-eating shark), derived from karcharos (= jagged) in reference to a shark’s rasp-like skin.
Squalus carcharias Linnaeus, 1758, Systema Naturae: 235. Type locality: Europe.
Bray, D.J. 2020
White Shark, Carcharodon carcharias (Linnaeus 1758)
Andreotti, S., von der Heyden, S., Henriques, R., Rutzen, M., Meÿer, M., Oosthuizen, H. & Matthee, C.A. 2015. New insights into the evolutionary history of white sharks, Carcharodon carcharias. Journal of Biogeography. doi: 10.1111/jbi.12641 Abstract
Blower, D.C., Pandolfi, J.M., Bruce, B.D., Gomez-Cabrera, M.C. & Ovenden, J.R. 2012. Population genetics of Australian white sharks reveals fine-scale spatial structure, transoceanic dispersal events and low effective population sizes. Marine Ecology Progress Series 455: 229–244.
Bonfil, R., M.P. Francis, C. Duffy, M.J. Manning & S.M. O’Brien. 2010. Large-scale tropical movements and diving behaviour of white sharks (Carcharodon carcharias) tagged off New Zealand. Aquatic Biology 8: 115–123.
Bonfil, R., Meer, M., Scholl, M.C., Johnson, R., O’Brien, S., Oosthuizen, H., Swanson, S., Kotze, D. & Paterson, M. 2005. Transoceanic migration, spatial dynamics, and population linkages of white sharks. Science 310(5745): 100-103. DOI: 10.1126/science.1114898.full Open access
Boustany, A.M., Davis, S.F., Pyle, P., Anderson, S.D., Le Boeuf B.J. & Block, B.A. 2002. Expanded niche for white sharks. Nature 415: 35–36.
Bradford, R. & Robbins, R. 2013. A rapid assessment technique to assist management of the White Shark (Carcharodon carcharias) cage dive industry, South Australia. The Open Fish Science Journal 6: 13-18.
Briscoe, D.A., Stow, A. & Hussey, S. 2006. Trial DNA Testing of shark products imported into and exported from Australia to detect the presence of Great White Shark. Report to the Department of the Environment and Heritage. May 2005.
Bruce, B.D. 1992. Preliminary observations on the biology of the White Shark, Carcharodon carcharias, in South Australian waters. Australian Journal of Marine and Freshwater Research 43: 1-11.
Bruce, B.D. 1995. The Protection of White Shark. A research perspective. Southern Fisheries 3(2): 11-15.
Bruce, B.D. 2006. Coastal migrations of white sharks in Australian waters: Evidence for site fidelity, temporary residency and common pathways (pp 67-77). In DPI NSW. Report from the Scientific Shark Protection Summit, 10 April 2006, 104 pp. PDF
Bruce, B.D. 2008. The Biology and Ecology of the White Shark, Carcharodon carcharias. In: Camhi, M.D, E.K. Pikitch & E.A Babcock (eds). Sharks of the Open Ocean. Pp 69-76. Oxford, UK: Blackwell Publishing.
Bruce, B.D & Bradford, R.W. 2008. Spatial dynamics & habitat preferences of juvenile white sharks: identifying critical habitat and options for monitoring recruitment. Final Report to the Department of the Environment, Water, Heritage and the Arts - Marine Species Recovery Program. Hobart: CSIRO.
Bruce, B. & Bradford, R.W. 2011. The effects of burleying on the distribution and behaviour of white sharks, Carcharodon carcharias, at the Neptune Islands, South Australia: final report. Department of Environment and Natural Resources, South Australia.
Bruce, B.D. & Bradford, R.W. 2012. Habitat use and spatial dynamics of juvenile white sharks, Carcharodon carcharias, in Eastern Australia. In Domeier, M. (ed.) Global perspectives on the biology and life history of the white shark. CRC Press, Boca Raton, FL.
Bruce, B.D. & Bradford, R.W. 2013. The effects of shark-cage diving operations on the behavior and movements of white sharks, Carcharodon carcharias, at the Neptune Islands, South Australia. Mar Biol ; DOI: 10.1007/s00227-012-2142-z
Bruce, B.D. & Bradford, R.W. 2015. Segregation or aggregation? Sex-specific patterns in the seasonal occurrence of white sharks Carcharodon carcharias, at the Neptune Islands, South Australia. Journal of Fish Biology 87: 1355-1370.
Bruce. B.D., Malcolm H. & Stevens J.D. 2001. A Review of the Biology and Status of White Sharks in Australian Waters. CSIRO Marine Research, Hobart.
Bruce, B.D., J.D. Stevens & R.W. Bradford. 2005. Site Fidelity, Residence Times and Home Range Patterns of White Sharks around Pinniped Colonies. [Online]. Final Report to the Australian Government Department of the Environment and Heritage, Canberra.
Bruce, B.D., Stevens, J.D. & Bradford, R.W. 2005. Identifying movements and habitats of white sharks and grey nurse sharks. [Online]. Available from: http://www.environment.gov.au/coasts/publications/white-grey-nurse-habitats/index.html.
Bruce, B.D., Stevens, J.D. & Malcolm, H. 2006. Movements and swimming behaviour of white sharks (Carcharodon carcharias) in Australian waters. Marine Biology 150: 161-172.
Cailliet, G.M., Natanson, L.J., Weldon, B.A. & Ebert, D.A. 1985. Preliminary studies on the age and growth of the white shark Carcharodon carcharias, using vertebral bands. Southern California Academy of Sciences, Memoirs 9: 49-60.
Carey, F.G., Kanwisher, J.W., Brazier, O., Gabrielson, G., Casey, J.G. & Pratt Jr, H.L. 1982. Temperature and activities of a white shark, Carcharodon carcharias. Copeia 1982: 254-260.
Castro, J. [Domeier, M.] 2012. A Summary of Observations on the Maximum Size Attained by the White Shark, Carcharodon carcharias. Global Perspectives on the Biology and Life History of the White Shark. Boca Raton, FL: CRC Press. pp. 85-90.
CITES (2004). Convention of International Trade in Endangered Species of Wild Fauna and Flora - Appendix II Listing of the White Shark (revision 1).
Compagno, L.J.V. 1984. FAO Species Catalogue. Sharks of the World. An annotated and illustrated catalogue of shark species known to date. Hexanchiformes to Lamniformes. FAO Fisheries Synopsis No. 125. Rome : FAO Vol. 4(1) pp. 1-249.
Compagno, L.J.V. 1998. Families Pseudocarchariidae, Alopiidae, Lamnidae. pp. 1268-1278 in Carpenter, K.E. & Niem, V.H. (eds). The Living Marine Resources of the Western Central Pacific. FAO Species Identification Guide for Fisheries Purposes. Rome : FAO Vol. 2 687-1396 pp.
Compagno, L.J.V. 2001. Sharks of the World. An annotated and illustrated catalogue of shark species known to date. Bullhead, mackerel and carpet sharks (Heterodontiformes, Lamniformes and Orectolobiformes). Rome : FAO, FAO Species Catalogue for Fisheries Purposes No. 1 Vol. 2 269 pp.
Compagno, L.J.V., Dando, M. & Fowler, S. 2005. A Field Guide to the Sharks of the World. London : Collins 368 pp.
Compagno, L.J.V., Marks, M.A. & Fergusson, I.K. 1997. Threatened fishes of the world: Carcharodon carcharias (Linnaeus, 1758) (Lamnidae). Environmental Biology of Fishes 50: 61-62.
Del Raye, G., Jorgensen, S.J., Krumhans, K., Ezcurra, J.M. & Block, B.A. 2013. Travelling light: white sharks (Carcharodon carcharias) rely on body lipid stores to power ocean-basin scale migration. Proc. R. Soc. B 280: 1471-2954. PDF
Department of the Enviroment, Water, Heritage and the Arts (DEWHA). 2009. White Shark Issues Paper. [Online]. Canberra, ACT: Australian Government.
Department of Sustainability, Environment, Water, Population and Communities. 2013. Recovery Plan for the White Shark (Carcharodon carcharias). Department of Sustainability, Environment, Water, Population and Communities. Available from: http://www.environment.gov.au/biodiversity/threatened/recovery-plans/recovery-plan-white-shark-carcharodon-carcharias. In effect under the EPBC Act from 06-Aug-2013 as Carcharodon carcharias.
Dicken, M.L. 2008. First observations of young of the year and juvenile great white sharks (Carcharodon carcharias) scavenging from a whale carcass. Marine and Freshwater Research 59: 596-602.
Domeier, M.L. 2012. A new life-history hypothesis for white sharks, Carcharodon carcharias, in the northeast Paciﬁc. Pp 199–223, in: Domeier, M. (ed.) Global perspectives on the biology and life history of the Great White Shark. CRC Press, Boca Raton.
Ellis, R. & McCosker, J.E. 1991. Great White Shark. Stanford University Press. 270 pp.
Estrada, J.A., Rice, A.N., Natanson, L J. & Skomal, G.B. 2006. Use of isotopic analysis of vertebrae in reconstructing ontogenetic feeding ecology in white sharks. Ecology 87: 829-834.
Fergusson, I., Compagno, L.J.V. & Marks, M.A. 2000. Predation by white sharks Carcharodon carcharias upon chelonians, with new records from the Mediterranean Sea and a first record of the ocean sunfish Mola mola as stomach contents. Environmental Biology of Fishes 58: 447-453.
Fergusson, I., Compagno, L.J.V. & Marks, M. 2009. Carcharodon carcharias. The IUCN Red List of Threatened Species 2009: e.T3855A10133872. http://dx.doi.org/10.2305/IUCN.UK.2009-2.RLTS.T3855A10133872.en. Downloaded on 02 October 2016.
Fitzpatrick, B., M. Meekan & A. Richards. 2006. Shark attacks on a whale shark (Rhincodon typus) at Ningaloo Reef, Western Australia. Bull. Mar. Sci. 78: 397-402.
Fowler, S.L., Cavanagh, R.D., Camhi, M., Burgess, G.H., Cailliet, G.M., Fordham, S.V., Simpfendorfer, C.A. and Musick, J.A. (comps and eds) 2005. Sharks, Rays and Chimaeras: The Status of the Chondrichthyan Fishes. Status Survey. pp. x + 461. IUCN/SSC Shark Specialist Group, IUCN, Gland, Switzerland and Cambridge, UK.
Francis, M.P. 1996. Observations on a pregnant White Shark with a review of reproductive biology (pp 157-172). In: Klimley, A.P. & D.G Ainley (eds). Great White Sharks. The biology of Carcharodon carcharias. Academic Press, San Diego.
Goldman, K.J. 1997. Regulation of body temperature in the white shark, Carcharodon carcharias. Journal of Comparative Physiology B. Biochemical Systemic and Environmental Physiology. 167: 423-429.
Goldman, K.J., Anderson, S.D., McCosker, J.E. & Klimley, A.P. 1996. Temperature, swimming depth, and movements of a White Shark at the South Farallon Islands, California. In: Klimley, A.P. & D.G Ainley (eds). Great White Sharks. The biology of Carcharodon carcharias. Academic Press, San Diego.
Grainger, R., Peddemors, V.M., Raubenheimer, D. & Machovsky-Capuska, G.E. 2020. Diet Composition and Nutritional Niche Breadth Variability in Juvenile White Sharks (Carcharodon carcharias). Frontiers in Marine Science https://doi.org/10.3389/fmars.2020.00422
Gubili, C., Duffy, C.A.J., Cliff, G., Wintner, S.P., Shivji, M., Chapman, D., Bruce, B.D., Martin, A.P., Sims, D.W. [Domeier M]. 2012. Application of molecular genetics for conservation of the White Shark, Carcharodon carcharias, L. 1758. Global Perspectives of the Life History of the White Shark. Boca Raton, FL: CRC Press. pp. 357-380.
Hammerschlag, N., Gallagher, A.J., Wester, J., Luo, J. & Ault, J.S. 2012. Don’t bite the hand that feeds: assessing the ecological impacts of provisioning ecotourism on an apex marine predator. Funct Ecol. doi:10.1111/j.1365-2435.2012.01973.x
Hutchins, B. & Swainston, R. 1986. Sea Fishes of Southern Australia. Complete Field Guide for Anglers and Divers. Swainston Publishing. 180 pp.
Johnson, J.W. 2010. Fishes of the Moreton Bay Marine Park and adjacent continental shelf waters, Queensland, Australia. 299-353 in Davie, P.J.F. & Phillips, J.A. Proceedings of the Thirteenth International Marine Biological Workshop, The Marine Fauna and Flora of Moreton Bay. Memoirs of the Queensland Museum 54(3) 
Jorgensen, S.J., C.A. Reeb, T.K. Chapple, S. Anderson, C. Perle, S.R. Van Sommeran, C. Fritz-Cope, A.C. Brown, A.P. Klimley & B.A. Block. 2010. Philopatry and migration of Pacific white sharks. Proc. R. Soc. B 277: 679–688. doi:10.1098/rspb.2009.1155 PDF
Klimley, A.P. 1994. The predatory behaviour of the White Shark. American Scientist 82: 122-133.
Klimley, A.P. & Ainley, D.G. (eds). 1996. Great White Sharks. The biology of Carcharodon carcharias. Academic Press, San Diego. 517 pp.
Klimley, A.P., Le Boeuf, B.J., Cantara, K.M., Richert, J.E., Davis, S.F., van Sommeran, S. & Kelly, J.T. 2001. The hunting strategy of white sharks (Carcharodon carcharias) near a seal colony. Marine Biology 138: 617-636.
Kuiter, R.H. 1993. Coastal Fishes of South-eastern Australia. Bathurst : Crawford House Press 437 pp.
Lack, M. & Sant, G. 2011. The Future of Sharks: A Review of Action and Inaction. Cambridge, U.K. and Washington D.C. TRAFFIC International and the Pew Environment Group. 45 pp.
Laroche, R.K., Kock, A.A., Dill, L.M. & Oosthuizen, W.H. 2007. Effects of provisioning ecotourism activity on the behaviour of white sharks, Carcharodon carcharias. Mar Ecol Prog Ser 338: 199-209.
Last, P.R. & Stevens, J.D. 1994. Sharks and Rays of Australia. Canberra : CSIRO Australia 513 pp. 84 pls
Last, P.R. & Stevens, J.D. 2009. Sharks and Rays of Australia. Vol 2. Collingwood : CSIRO Publishing Australia, 656pp.
Lingham-Soliar, T. 2005. Caudal fin allometry in the white shark Carcharodon carcharias: implications for locomotory performance and ecology. Die Naturwiss. 92: 231– 236. (doi:10.1007/s00114-005-0614-4)
Linnaeus, C. 1758. Systema Naturae per Regna tria Naturae, secundem Classes, Ordines, Genera, Species, cum Characteribus, Differentis, Synonymis, Locis. Tom.1, Editio decima, reformata. Holmiae : Laurentii Salvii 824 pp.
Macbeth, W.G., Vandenberg, M. & Graham, K.J. 2008. Identifying Sharks and Rays; a Guide for Commercial Fishers. Sydney : New South Wales Department of Primary Industry 71 pp.
Malcolm, H., Bruce, B.D. & Stevens, J. 2001. A Review of the Biology and Status of White Sharks in Australian Waters. Environment Australia, Canberra. 113 pp.
Mcauley, R.D., Bruce, B.D., Keay, I.S., Mountford, S., Pinnell, T. & Whoriskey, F.G. 2017. Broad-scale coastal movements of white sharks off Western Australia described by passive acoustic telemetry data. Marine and Freshwater Research. 68(8) 1518-1531. https://doi.org/10.1071/MF16222 Abstract
McCoy, F. 1883. Carcharodon rondeletii (Mul. & Hen.) The White Shark, Pl. 74, pp. 19-22, in McCoy, F. The Prodromus of the zoology of Victoria; figures and descriptions of the living species of all classes of the Victoria. Decade VIII: 1-36, Pls 71-80. See ref at BHL
McPherson, J.M & Myers, R.A. 2009. How to infer population trends in sparse data: examples with opportunistic sighting records for great white sharks. Diversity and Distributions. 15: 880-890.
Müller, J. & Henle, F.G.J. 1839. Systematische Beschreibung der Plagiostomen. Berlin : Veit & Co pp. 29-102 pls. (described as Carcharodon rondeletii) See ref at BHL
Pardini, A.T., Jones, C.S., Noble, L.R., Kreiser, B., Malcolm, H., Bruce, B.D., Stevens, J.D., Cliff, G., Scholl, M.C., Francis, M., Duffy, C. & Martin, A.P. 2001. Philopatric females and roving male great white sharks. Nature 412: 139–140.
Pardini, A.T., Jones, C.S., Noble, L.R., Kreiser, B., Malcolm, H., Bruce, B.D., Stevens, J.D., Cliff, G., Scholl, M.C., Francis, M., Duffy, C. & Martin, A.P. 2001. Sex biased dispersal of Great White Sharks. Nature 412: 139-140.
Pepperell, J. 1992. Trends in the distribution, species composition and size of sharks caught by gamefish anglers off south-eastern Australia, 1961-1990. In: Pepperell, J. (ed.) Sharks: Biology and Fisheries. Australian Journal of Marine and Freshwater Research, pp. 213-225.
Pepperell, J. 2010. Fishes of the Open Ocean a Natural History & Illustrated Guide. Sydney : University of New South Wales Press Ltd 266 pp.
Pratt, Jr. H.L. 1996. Reproduction in the male White Shark. In: Klimley, A.P. & Ainley, D.G. (eds). Great White Sharks: The Biology of Carcharodon carcharia. Academic Press, San Diego.
Randall, J.E. 1973. Size of the great white shark (Carcharodon). Science (Washington, D.C.) 181(4095): 169-170
Reid, D.D. & Krogh, M. 1992. Assessment of catches from Protective Shark meshing off New South Wales beaches between 1950 and 1990. Australian Journal of Marine and Freshwater Research 43(1): 283-296.
Robbins, R.L. 2007. Environmental variables affecting the sexual segregation of great white sharks Carcharodon carcharias at the Neptune Islands South Australia. Journal of Fish Biology 70: 1350-1364.
Robbins, R.L. & Booth, D.J. 2012. Seasonal sexual and size segregation of white sharks, Carcharodon carcharias, at the Neptune Islands, South Australia. Pp. 287-99, in Domeier, M.L. (ed.) Global perspectives on the biology and life history of the white shark. Boca Raton, Fl: CRC Press.
Robbins, R.L., Enarson, M., Bradford, R.W., Robbins, W.D. & Fox, A.G. 2015. Residency and Local Connectivity of White Sharks at Liguanea Island: A Second Aggregation Site in South Australia? The Open Fish Science Journal 8: 23-29. [DOI: 10.2174/1874401X01508010023] PDF Open Access
Saïdi, B., Bradaï, M.N., Bouaïn, A., Guélorget, O. & C. Capapé. 2005. Capture of a pregnant female white shark, Carcharodon carcharias (Lamnidae) in the Gulf of Gabés (southern Tunisia, central Mediterranean) with comments on oophagy in sharks. Cybium 29(3): 303-307.
Sato, K., Nakamura, M., Tomita, T., Toda, M., Miyamoto, K. & Nozu, R. 2016, How great white sharks nourish their embryos to a large size: evidence of lipid histotrophy in lamnoid shark reproduction. Biology Open 5: 1211-1215; doi: 10.1242/bio.017939 Open access
Schilds, A., Mourier, J., Huveneers, C., Nazimi, L., Fox, A. & Leu, S.T. 2019. Behavioral Ecology and Sociobiology 73(10): 138. https://doi.org/10.1007/s00265-019-2745-1
Semmens, J.M., Payne, N.L., Huveneers, C., Sims, D.W. & Bruce, B.D. 2013. Feeding requirements of white sharks may be higher than originally thought. Scientific Reports 3: 1471
Shivji, M.S., Chapman, D.D., Pikitch, E.K. & Raymond, P.W. 2005. Genetic profiling illegal international trade in fins of the great white shark, Carcharodon carcharias. Conservation Genetics 6: 1035-1039.
Sims, D.W., Humphries, N.E., Bradford, R.W. & Bruce, B.D. 2012. Levy ﬂight and brownian search patterns of white sharks in response to different prey ﬁelds. J Anim Ecol 81: 432–442.
Stevens, J.D. 1994. Family Lamnidae. pp. 141-143 figs 112-114 in Gomon, M.F., Glover, C.J.M. & Kuiter, R.H (eds). The Fishes of Australia's South Coast. Adelaide : State Printer 992 pp. 810 figs
Strong, W.R., Nelson, D.R., Bruce, B.D. and Murphy, R.C. 1996. Population dynamics of White Sharks in Spencer Gulf, South Australia (pp. 401-414). In: Klimley, A.P. & D.G. Ainley, (eds). Great White Sharks - The Biology of Carcharodon carcharias. Academic Press, San Diego.
Strong, W.R., Murphy, R.C., Bruce, B.D. and Nelson, D.R. 1992, Movements and associated observations of bait-attracted White Sharks, Carcharodon carcharias: A preliminary report. Australian Journal of Marine and Freshwater Research 43: 13-20.
Tricas, T.C. & McCosker, J.E. 1984. Predatory behaviour of the white shark (Carcharodon carcharias) with notes on its biology. Proceedings of the California Academy of Sciences. 43: 221-238.
Uchida, S., Toda, M., Teshima, K. and Yano, K. 1996. Pregnant White Shark and full-term embryos from Japan, pp 139-155, in Klimley, A.P. & Ainley, D.G. (eds). Great White Sharks - The Biology of Carcharodon carcharias. Academic Press, San Diego.
Walker, T. 1999. Protection for Great Whites. Sport Fishing Issue 5.
White, W. 2008. Shark Families Heterodontidae to Pristiophoridae. pp. 32-100 in Gomon. M.F., Bray, D.J. & Kuiter, R.H (eds). Fishes of Australia's Southern Coast. Sydney : Reed New Holland 928 pp.
Whitley, G.P. 1934. Notes on some Australian sharks. Memoirs of the Queensland Museum 10(4): 180-200 figs 1-4 pls 27-29 (as Carcharhinus carcharias)
Whitley, G.P. 1939. Taxonomic notes on sharks and rays. The Australian Zoologist 9(3): 227-262 figs 1-18 pls 20-22 (described as Carcharodon albimors)
Whitley, G.P. 1940. The Fishes of Australia. Part 1. The sharks, rays, devil-fish, and other primitive fishes of Australia and New Zealand. Sydney : Roy. Zool. Soc. N.S.W. 280 pp. 303 figs. (as Carcharodon albimors)
Whitley, G.P. 1967. Sharks of the Australasian region. The Australian Zoologist 14(2): 173-188 fig. 1 pl. 4 (as Carcharodon albimors)