Velvet Dogfish, Zameus squamulosus (Günther 1877)

A Velvet Dogfish, Zameus squamulosus, from the abyssal plain between the Norfolk Ridge and Lord Howe Rise, Tasman Sea - depth 1635-1749 m. Source: Robin McPhee & Mark McGrouther / NORFANZ Founding Parties. License: All rights reserved

A black bioluminescent shark with small dorsal fin spines, a moderately long snout, small lanceolate teeth without cusplets in the upper jaw, and large high, knife-cusped cutting teeth in the lower jaw, and the caudal fin with a strong subterminal notch and a short lower lobe.

Cite this page as:
Dianne J. Bray, Zameus squamulosus in Fishes of Australia, accessed 02 Mar 2024,

Velvet Dogfish, Zameus squamulosus (Günther 1877)

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Off Western Australia from Cape Naturaliste to Rowley Shoals, and off northern New South Wales to off Tasmania. Elsewhere, the species is widespread but found sporadically in most oceans except the Eastern Pacific. 
Velvet Dogfish are pelagic or occur on or near the bottom, near continental slopes and seamounts.


Aplacental viviparous (ovoviviparous).

Species Citation

Centrophorus squamulosus Günther, 1877, Ann. Mag. Nat. Hist. 4 20(56): 433. Type locality: off Inosima, Japan, ♂.


Dianne J. Bray

Velvet Dogfish, Zameus squamulosus (Günther 1877)


Burgess, G.H. & Chin, A. 2006. Zameus squamulosus. The IUCN Red List of Threatened Species. Version 2015.2. . Downloaded on 19 August 2015.

Compagno, L.J.V. 1984. FAO Species Catalogue. Sharks of the World. An annotated and illustrated catalogue of shark species known to date. Hexanchiformes to Lamniformes. FAO Fisheries Synopsis No. 125. Rome : FAO Vol. 4(1) pp. 1-249.

Compagno, L.J.V., Dando, M. & Fowler, S. 2005. A Field Guide to the Sharks of the World. London : Collins 368 pp.

Compagno, L.J.V. & Niem, V.H. 1998. Hexanchidae, Echinorhinidae, Squalidae. pp. 1208-1232 in Carpenter, K.E. & Niem, V.H. (eds). The Living Marine Resources of the Western Central Pacific. FAO Species Identification Guide for Fisheries Purposes. Rome : FAO Vol. 2 687-1396 pp.

Daley, R.K., Stevens, J.D., Last, P.R. & Yearsley, G.K. 2002. Field Guide to Australian Sharks & Rays. Hobart : CSIRO Marine Research 84 pp.

Günther, A. 1877. Preliminary notes on new fishes collected in Japan during the expedition of H.M.S. Challenger. Annals and Magazine of Natural History 4 20(56): 433-446.

Jordan, D.S. & Fowler, H.W. 1903. A review of the elasmobranchiate fishes of Japan. Proceedings of the United States National Museum 26(1324): 593-674.

Last, P.R. & Stevens, J.D. 1994. Sharks and Rays of Australia. Canberra : CSIRO Australia 513 pp. 84 pls.

Last, P.R. & Stevens, J.D. 2009. Sharks and Rays of Australia. Collingwood : CSIRO Publishing Australia 2, 550 pp.

Regan, C.T. 1908. A synopsis of sharks of the family Squalidae. Annals and Magazine of Natural History 8 2: 39-57.

Quick Facts

CAAB Code:37020042


Conservation:IUCN Data Deficient

Depth:550-2000 m (usually)

Max Size:69 cm TL

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Species Maps

CAAB distribution map